A post-doctoral position is opened at the Institut des Sciences de l?Evolution, (CNRS-Universit? Montpellier), Montpellier, France, and at the Station Biologique de Roscoff (CNRS-Universit? Paris 6), Brittany, France.

The proposed work consists in the development of theoretical population genetics models to study genetic hitchhiking in spatially structured populations. Recent methods use measures of genetic differentiation (Fst) at neutral loci in order to locate genes that are selected differently in different regions of a species distribution range (?genome scans?). However, the theory on the effects of selection on linked, neutral variation in spatially structured populations remains little developed. A combination of mathematical and simulation approaches will be used to explore the effects of different adaptive/demographic scenarios on patterns of neutral variation (allele frequency spectra, linkage disequilibrium...) in spatially structured populations. These models will be used to identify whether some statistics (if not at the level of a single locus, at the level of a whole scan) may be used to discriminate between these scenarios. Genome scan data from different species of marine invertebrates will be available in order to test these methods. Once this first work accomplished, a second subject will be to develop models of speciation in spatially structured populations, focusing on the interaction between heterogeneous selection and the evolution of endogeneous (Dobzhansky-Muller) incompatibilities.

Candidates should have a strong background in theoretical population genetics. Strong motivation, communication skills and commitment are expected. Although knowledge of French is helpful in every day life, the working language will be English.

The position starts with a first period of 14 months at the Institut des Sciences de l'Evolution (Montpellier-S?te, France, http://www.isem.cnrs.fr), starting between April and September 2009, and will be expandable to a second period of 14 months at the Station Biologique de Roscoff (Brittany, France, http://www.sb-roscoff.fr). The salary will range from 1800 ? to 2500? per month depending on the experience of the candidate.

Applicants should send a CV including summary of past research, current interests and motivation for the present project, as well as contact information for three references to Denis Roze (roze@sb-roscoff.fr) and Nicolas Bierne (n-bierne@univ-montp2.fr).

Denis Roze

Adaptation et Diversit? en Milieu Marin

Station Biologique de Roscoff

Place Georges Teissier, BP 74

29682 Roscoff Cedex, France

Ph: (+33) 2 98 29 23 20

Fax: (+33) 2 98 29 23 36

Nicolas Bierne

D?partement "Biologie Int?grative"

Institut des Sciences de l'Evolution

UMR5554 CNRS - Universit? de Montpellier 2 Station M?diterran?enne l'Environnement Littoral 1,

Quai de la Daurade

34200 S?te FRANCE T?l?

phone : +33 (0)4 67 46 33 75 T?l?

copie : +33 (0)4 67 46 33 99

E-mail. n-bierne@univ-montp2.fr http://www.isem.cnrs.fr/spip.php?rubrique139 http://kimura.univ-montp2.fr/~nbierne/NicolasBierneCV.htm http://kimura.univ-montp2.fr/~nbierne/GenePhys.html